As I'm sure you've noticed reading this article,
The volume of a viral capsid is important not only because of the size of the viral genome, but also because nucleic acids carry a self-repulsing negative charge that must be physically contained. These repeating negative charges are distributed along the length of the nucleic acid molecules, and in solution, they are surrounded by positively charged ions like Na+ and Mg2+ or by more complex molecules like polyamines, which screen the negative charge of the phosphate backbone.
，这一点在搜狗输入法中也有详细论述
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This tells Coalton that two Integer values will be returned. The lisp operator now always needs the -, even if just a single value is returned.